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Retrieved Human Allografts A Clinicopathological Study
William F. Enneking, MD; Domenico A. Campanacci, MD
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Investigation performed at the Department of Orthopaedics and Rehabilitation, University of Florida College of Medicine, Gainesville, Florida
William F. Enneking, MD
Department of Orthopaedics and Rehabilitation, University of Florida College of Medicine, Box 100246 JHM Health Center, Gainesville, FL 32610-0246

Domenico A. Campanacci, MD
First University Clinic, Instituto Ortopedico Rizzoli, Via Pupilli 1, 40100 Bologna, Italy

No benefits in any form have been received or will be received from a commercial party related directly or indirectly to the subject of this article. No funds were received in support of this study.

The Journal of Bone & Joint Surgery.  2001; 83:971-986 
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Abstract

Background: We studied seventy-three massive preserved human allografts, retrieved from two to 156 months after implantation, to provide insight into the mechanisms of their repair.

Methods: The specimens were studied with radiographic and histological techniques that permitted time-related quantitative analysis of the reparative mechanisms of union, cortical repair, soft-tissue attachment, fracture, and characteristics of the allograft-cement interface and the articular cartilage.

Results: Union at cortical-cortical junctions occurred slowly (approximately twelve months) by host-derived external callus that bridged the junction and filled the gap between abutting cortices. The bone in the gap did not undergo stress-oriented remodeling even after many years, and, when the union was intentionally disrupted, failure occurred at the cement line that marked the allograft-host junction. Repair of the necrotic graft matrix was both external and internal. External repair consisted of the apposition of a thin seam of host bone on the outer surface of the graft, coating about 40% of the surface at one year and 80% at two years. Internal repair was confined to the ends and the periphery of the cortices and penetrated so slowly that only 15% to 20% of the graft was repaired by five years, after which deeper repair seldom occurred. Graft fractures in specimens retrieved soon after fracture showed only necrotic bone adjacent to the fracture site, whereas those retrieved after fracture-healing showed a marked increase in internal repair of the bone about the fracture site. When bone cement had been used to fix a prosthesis, there was no evidence of bone resorption or loosening of the device.

The osteoarticular specimens showed no survival of chondrocytes in the articular cartilage. However, the architecture of the acellular cartilage was well preserved after two to three years and occasionally after as many as five years. Late degenerative changes in the articular cartilage coincided with subchondral revascularization and fragmentation, and the articulating surfaces became covered by a pannus of fibrovascular reparative tissue. Degenerative changes in articular cartilage occurred earlier and were more advanced in specimens retrieved from patients with an unstable joint than in those retrieved from patients with a stable joint.

Conclusions: Repair of massive human allografts is an indolent process that follows a fairly predictable course during the first few years and is influenced by other biological activities, such as fracture repair, supplementary autografting, and tumor recurrence.

Clinical Relevance: These observations provide a clear, detailed picture of the extent, timing, and deficiencies in the incorporation and repair of large human allografts preserved by conventional banking techniques. As such, they provide a basis for comparative studies of the efficacy of the recently developed osteoinductive substances currently under investigation.

Figures in this Article
    In 1991, we described the radiographic and histological findings in sixteen massive retrieved human allografts that had been in situ from four to sixty-five months1. In the ensuing eight years, we studied an additional fifty-seven specimens that had been in situ from two to 156 months. The findings from these additional specimens promote a more complete understanding of the characteristics and deficiencies of the repair of human allografts and their clinical importance.
     
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    +Fig. 1:Graph showing the number of specimens retrieved per year.
     
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    +Fig. 2-A:Figs. 2-A and 2-B The histological features of cortical-cortical union. Fig. 2-A A low-power photomicrograph showing the junction between the acellular cortex of an allograft (AG) and the host bone (HB) filling the gap (hematoxylin and eosin, 20). The allograft was retrieved at five years. The site of union is marked by a distinct cement line (CL). Only a single haversian canal (HC) has been revascularized.
     
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    +Fig. 2-B:Figs. 2-A and 2-B The histological features of cortical-cortical union. Fig. 2-B A low-power photomicrograph showing the junction between the cortex of an allograft (AG) and the host bone (HB) filling the gap (hematoxylin and eosin, 20). The allograft was retrieved at eleven years. The site of union is still marked by a distinct cement line (CL). Revascularization has penetrated two haversian canals (HC).
     
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    +Fig. 3-A:Figs. 3-A and 3-B The site of failure at forty-two months. Fig. 3-A A photomicrograph showing the failure site between the cortex of an allograft (AG) and the host cortex (HE), which has extended through the mature external callus (EC) that united the two cortices (hematoxylin and eosin, 4).
     
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    +Fig. 3-B:Figs. 3-A and 3-B The site of failure at forty-two months. Fig. 3-B A photomicrograph showing the failure site between the allograft (AG) and the nonoriented host bone (HB) at the demarcating cement line (CL) (hematoxylin and eosin, 40).
     
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    +Fig. 4-A:Figs. 4-A and 4-B Union by endosteal callus. Fig. 4-A Anteroposterior radiograph of a proximal tibial osteoarticular allograft immediately before retrieval at seventeen months. No external callus is visible, and there is no radiolucent line at the junction (J) of the allograft and the more osteopenic host tibia.
     
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    +Fig. 4-B:A low-power photomicrograph of the allograft cortices (AG) and the more osteopenic host cortices (HC) (hematoxylin and eosin, 10). The upper junction is united by internal endosteal callus (IC), while the lower junction remains ununited despite the well-formed internal callus (IC).
     
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    +Fig. 5:A photomicrograph of the surface of an allograft retrieved eleven months after implantation, showing surface repair of the allograft cortical bone (hematoxylin and eosin, 120). The host soft tissues (HST) are apposed to the surface of the allograft cortex. A bud of reparative tissue entered the surface stoma of a haversian canal (HC), enlarging it by osteoclastic resorption. A thin seam of bone (HB) has been laid down on the surface of the allograft and is demarcated from it by a distinct cement line (CL).
     
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    +Fig. 6-A:Figs. 6-A and 6-B Revascularization at the osteotomy site. Fig. 6-A A photomicrograph showing the cortical osteotomy site of an allograft retrieved nineteen months after implantation (hematoxylin and eosin, ¥60). The three arrows in the center of the figure indicate the site of union between the host bone (HB) and the allograft (AG). Reparative tissue has penetrated into the graft in two areas (RT), enlarging the haversian canals (HC), but several canals show no evidence of repair.
     
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    +Fig. 6-B:A photomicrograph showing the cortical osteotomy site of an allograft retrieved twenty-five months after implantation (hematoxylin and eosin, ¥90). The viable host bone (HB) is united to the necrotic allograft (AG). The six arrows at the left of the figure indicate a previously enlarged haversian canal, filled with repair bone (RB). A second haversian canal contains reparative tissue (RT). Other haversian canals (HC) remain unrepaired.
     
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    +Fig. 7-A:Figs. 7-A and 7-B Internal repair. Fig. 7-A A low-power photomicrograph of a repairing allograft retrieved at thirty months (hematoxylin and eosin, 20). The host muscle (HM) is separated from the repairing allograft (AG) by a thin zone of proliferating host fibrovascular repair tissue. The haversian canals of the superficial osteons (SO) are largely excavated, leaving the interstitial lamellae (IL) unrepaired, while the deeper osteons are not yet revascularized and their haversian canals (HC) are not yet excavated.
     
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    +Fig. 7-B:A photomicrograph of osteons being reformed in a cortical allograft retrieved at twenty-six months (hematoxylin and eosin, 90). Osteoclastic resorption has almost reached the cement line (arrows) bounding the osteon on the left, while on the right excavation has reached the bounding cement line (arrows) about the osteon, the osteoclasts have disappeared, and osteoblastic rebuilding (OB) has begun.
     
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    +Fig. 8-A:Figs. 8-A through 8-E Reossification of allograft resorption. Fig. 8-A Anteroposterior radiograph made after implantation of the initial osteoarticular allograft.
     
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    +Fig. 8-B:Figs. 8-A through 8-E Reossification of allograft resorption. Fig. 8-B Anteroposterior radiograph made fourteen months later.
     
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    +Fig. 8-C:Fig. 8-C Anteroposterior radiograph made after implantation of the second osteoarticular allograft.
     
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    +Fig. 8-D:Fig. 8-D Anteroposterior radiograph made six months later.
     
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    +Fig. 8-E:Fig. 8-E Anteroposterior radiograph made twelve years after curettage and insertion of a supplementary vascular autogenous fibular graft.
     
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    +Fig. 9-A:Figs. 9-A through 9-D Union of an allograft-allograft junction. Fig. 9-A Postoperative anteroposterior radiograph of a 27-cm femoral osteoarticular allograft. The arrow indicates the allograft-host junction (J).
     
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    +Fig. 9-B:Figs. 9-A through 9-D Union of an allograft-allograft junction. Fig. 9-B Anteroposterior radiograph made four years postoperatively, at the time of distal allograft fracture. The allograft-host junction is well healed (arrow).
     
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    +Fig. 9-C:Fig. 9-C Anteroposterior radiograph made one year after a second reconstruction with a 15-cm allograft-prosthesis composite. The arrow indicates the junction of the first and second allografts.
     
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    +Fig. 9-D:Fig. 9-D Magnified anteroposterior radiograph of the junction of the first and second allografts, made seven years after the second reconstruction.
     
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    +Fig. 10:Early degenerative changes in articular cartilage. A photomicrograph of the articular surface of a proximal tibial osteoarticular allograft retrieved at eleven months (hematoxylin and eosin, 60). The cartilage is acellular and has severe degenerative changes, although the tidemark (TM) remains intact. The underlying subchondral marrow spaces have been revascularized, although there has been little repair of bone.
     
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    +Fig. 11:Pannus formation. A photomicrograph of the posterior aspect of the articulating surface of a femoral condyle from a distal femoral osteoarticular allograft retrieved at seventeen months (hematoxylin and eosin, 40). The articular cartilage (C) is acellular. A thick pannus of fibrovascular tissue (FVT) covers the cartilage. A tongue of more cellular tissue invades the cartilage at its periphery, having crossed both the tidemark (TM) and the osteochondral cement line (CL).
     
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    +Fig. 12-A:Figs. 12-A and 12-B Survival of fresh allograft chondrocytes. Fig. 12-A A photomicrograph of a fresh osteoarticular distal femoral allograft retrieved at thirteen months (hematoxylin and eosin, 5). The architecture of both the articular cartilage and the underlying femoral condyle is well preserved.
     
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    +Fig. 12-B:A photomicrograph of a representative field of the articular cartilage shows the normal architecture of adult articular cartilage (hematoxylin and eosin, 90). The vast majority of the chondrocytes appear viable, with only small patches of acellular matrix. There is no evidence of an inflammatory response.
    From 1975 to 1998, 405 preserved allografts were used to reconstruct skeletal defects created by limb-salvaging resection for the treatment of musculoskeletal tumors. Sixty specimens retrieved from these patients and thirteen other specimens provided by other institutions form the basis of these observations.

    Reasons for Retrieval

    Forty-two specimens (58%) were retrieved because of a complication associated with the allograft. Twenty-four specimens (33%) were retrieved because of a complication associated with a tumor: twelve were retrieved following amputation because of a local recurrence and twelve were retrieved at autopsy after death from pulmonary metastases. Seven other specimens were retrieved because of miscellaneous reasons.

    Types of Grafts

    Sixty-six (90%) of the specimens were massive allografts: thirty-seven were osteoarticular, twenty-one were intercalary, and eight were composite. Seven specimens (10%) were intercalary grafts that had been either inlaid or onlaid in a large defect: four were fibular grafts and three were corticocancellous iliac-crest grafts.

    Sites of Grafts

    Sixty-five massive grafts were used in reconstruction of the femur (forty-one), tibia (thirteen), or humerus (eleven), and eight were used in other sites.

    Duration in Situ (Fig. 1)

    The grafts were retrieved from two to 156 months after implantation. Fifty-four (74%) were retrieved during the first three years after implantation. The average duration in situ was twenty-eight months for those retrieved because of complications associated with the graft and thirty months for those retrieved because of complications associated with a tumor.

    Patient Demographics

    The specimens were retrieved from thirty-three male and forty female patients. The average age at the time of retrieval was twenty-eight years (range, five to sixty-nine years). Forty-three patients had received preoperative and/or postoperative chemotherapy, and three patients had received postoperative radiation therapy.

    Methods

    Various combinations of radiographic, macroscopic, and histological studies were done on the specimens, and these observations were correlated with the clinical and radiographic data, as described in our earlier study1.
    After the initial dissection, radiographs and photographs of the intact specimens were made and then the specimens were dissected. The cut surfaces of the bisected slabs were photographed under incandescent light and, after in vivo labeling with tetracycline had been done, under ultraviolet light.
    The slabs were fixed, decalcified, embedded in celloidin, cut into 20-m sections, and stained with hematoxylin and eosin. These macrosections were studied with low-power microscopy (with a magnification of four to ten times), and spatial maps were prepared from tracings of the macrosections to determine the extent and distribution of revascularization and repair. On these spatial maps, areas of host callus, appositional new-bone formation on the surface of the allograft, osteoclastic bone resorption, internal cortical revascularization, and osteonal remodeling were color-coded for area measurements by planimetry. These features were then converted to proportions of the specimen as quantitative estimates of the various reparative processes. Selected regions of the specimens were decalcified, embedded in paraffin, cut into 5-m sections, and stained with hematoxylin and eosin. When articular cartilage was examined, toluidine blue and safranin-O stains were also used. These specimens were studied with high-power microscopy.

    Union

    Two types of junctions were available for study. Most were cortical-cortical (forty-four specimens), whereas a few were cancellous-cancellous (twelve specimens). At ten of the cortical-cortical junctions, additional autogenous cancellous grafts had been onlaid.

    Cortical-Cortical Junctions

    At the cortical-cortical junctions, healing took place by bridging external callus that originated from the periosteum of the host bone and extended for up to 3 cm on the surface of the allograft, where it became annealed (Fig. 2-A). The callus did not appear to have extended from both sides of the junctions (host and graft) and then to have joined at the osteotomy site in any specimen. The gap at the osteotomy site initially filled with reparative fibrovascular tissue from the surrounding host soft tissues. This, in turn, was replaced by immature trabeculae that gradually matured into haversian bone. The external callus about the osteotomy site consistently was more mature than the bone filling the gap. Although immature trabeculae were observed in the gap in specimens retrieved at four months, maturation into haversian bone was not consistently seen until twelve months. Radiographs demonstrated a persistent radiolucent gap at the osteotomy site, even when it was filled with mature lamellar trabeculae joining the two cortices, until the uniting bone within the gap had converted to haversian bone. When the bone filling the gap had matured into cortical bone, the haversian canals in the uniting seam were oriented perpendicularly to the long axis of the bone, presumably because of the orientation of the initial ingrowth of blood vessels from the periphery of the gap. The site of union between the uniting seam and the abutting allograft remained clearly marked by a distinct cement line. Little or no fibrovascular repair tissue penetrated into the haversian canals of the allograft, even in specimens that had been in situ for up to five years. The nonstress orientation of the bone in the gap, however, persisted for years and was seen as late as eleven years without remodeling parallel to the long axis of the bone (Fig. 2-B).
    In two specimens (retrieved at twenty-eight and forty-two months) that were subjected to torque to failure at the junctions, the failure occurred exactly along the cement line that denoted the histological line of union (Figs. 3-A and 3-B). None of the specimens had primary union between abutting host and allograft cortices. Only one specimen had areas of endochondral ossification. Correlation between the radiographic and histological findings showed agreement in all but three specimens. These specimens, each in situ for less than a year, showed a persistent radiolucent gap at the osteotomy site. However, on histological examination, they demonstrated immature trabeculae uniting the gap.
    Three specimens that had been fixed with a plate demonstrated no meaningful external callus at the cortical-cortical junctions. Rather, a bridge of cancellous bone extended from the medullary canal of the host, filled the osteotomy site, and extended up the canal of the allograft for 2 to 3 cm (Figs. 4-A and 4-B).
    Seventeen cortical-cortical junctions in specimens from patients treated with preoperative and postoperative chemotherapy were available for histological evaluation. Of the nine specimens retrieved within twelve months, only one showed immature union. Of the eight specimens retrieved between fourteen and twenty-eight months, five showed immature union, two had haversian union, and one contained only mature fibrovascular reparative tissue in the gap.

    Cortical-Cortical Nonunions

    Six allografts were retrieved because of nonunion, and one was found to have nonunion when it was retrieved because of tumor recurrence. Two (mentioned above) were retrieved from patients who had received chemotherapy, and one (the graft that was retrieved because of local recurrence) was from a patient who had received radiation therapy. The remaining four were retrieved more than twelve months after implantation from patients who had not received chemotherapy or radiation therapy. All of these specimens had similar histological findings; a varying amount of external callus extended from the host cortex but was prevented from annealing to the allograft by an envelope of fibrovascular tissue filling the gap and encasing the end of the allograft. The tissue contained few inflammatory cells and occasional histiocytes and multinucleated giant cells. Gram stains and cultures of this tissue were negative. The surfaces of the abutting allograft were intermittently pockmarked with Howship lacunae, with some containing osteoclasts, which resulted in only modest amounts of resorption radiographically.
    Of three specimens that were retrieved because of infection and had been in situ for more than twelve months, two in which the infection involved the cortical-cortical junction did not have union. In these two specimens, the osteotomy gap was filled with chronic inflammatory tissue and there was intense osteoclastic resorption of the adjacent allograft. In the third specimen, a composite allograft-total hip replacement, the infection was confined to the periprosthetic tissues and the uninvolved distal junction was well healed.
    Of the ten specimens in which supplementary autogenous cancellous grafts had been placed about the cortical-cortical junction, eight had new-bone formation uniting the autograft to the surface of the allograft as early as six months after implantation. When the autogenous graft was in contact with the allograft, there was not only intense new-bone formation on the surface of the graft but also, surprisingly, more extensive internal repair in the underlying cortex of the allograft than in the remainder of the graft. In the two specimens in which there was no union between the autogenous graft and the allograft by either an external callus or a seam of bone between the abutting cortices, the autograft was separated by a zone of fibrovascular repair tissue from the external surface of the allograft. Similar observations were noted in the four specimens in which a vascularized autogenous fibular graft had been placed in juxtaposition to the allograft. In each specimen, the vascularized graft had united to the allograft and the region of the allograft beneath the site of union had been much more extensively repaired than the remainder of the allograft.

    Cancellous-Cancellous Junctions

    Union at the cancellous-cancellous junctions took place uniformly and was present as early as four months. These specimens had no external callus. Fibrovascular repair tissue from the host bone had invaded the marrow spaces of the allograft and deposited seams of reparative bone on the surfaces of the trabeculae of the allograft, uniting them to the trabeculae of the host bone. The combination of new bone encasing the preexisting allograft trabeculae produced substantially thickened trabeculae, which was reflected radiographically by a narrow zone of increased radiodensity just on the allograft side of the junction. In comparisons of the macrosections and the radiographs of the cancellous junctions, the 2 to 4-mm zone of radiodensity accurately portrayed the depth of penetration of trabecular repair. Deep to this zone of thickened, repaired trabeculae, the original architecture of the allograft was well preserved, but the marrow spaces were filled with dense hypovascular fibrous tissue that appeared to block additional penetration of the allograft by host reparative fibrovascular tissue. Beyond this zone, deeper in the allograft, the acellular trabeculae preserved the original architecture of the allograft, while the marrow spaces were filled with the acellular remnants of fatty marrow. Even after many years, the cancellous area deep to the fibrous barrier remained composed of undisturbed necrotic remnants of allograft marrow and intact but acellular trabeculae.

    Cortical Repair

    The extent and distribution of cortical revascularization and repair were studied in fifty-eight massive allografts. Repair of the cortices was both external and internal.

    First Year

    Sixteen specimens were retrieved at less than twelve months. The external surface of the allografts had scattered areas of osteoclastic resorption, which were particularly evident where there was close contact with host soft tissue. The osteoclastic resorption formed small erosive cavities that were filled with fibrovascular reparative tissue and occasionally were still occupied by an osteoclast. These cavities occupied, on average, approximately 20% of the external surface of the graft. In between the resorption cavities, a 1 to 2-mm seam of appositional new bone was annealed by a cement line to the unresorbed surface of the allograft. This newly deposited surface bone was seen as early as four months and by one year occupied an average of approximately 40% of the surface of the graft. Associated with these surface changes, buds of fibrovascular tissue penetrated into the Volkmann canals on the surface of the graft (Fig. 5). Occasionally, the penetrating fibrovascular buds were accompanied by osteoclasts enlarging the canals, and, where the canals had been enlarged, thin seams of new bone lined the canal for a distance of 1 to 2 mm. The penetration by the revascularizing buds from the external surface rarely exceeded 2 mm during the first year.
    At the osteotomy site, penetration of the transected haversian canals in the graft cortices by buds of fibrovascular repair tissue was rarely seen, and, when present, it extended for only 1 to 2 mm. Osteoclastic resorption of the edges of the revascularized canals was also unusual. Direct osteoclastic resorption into the exposed surface, a so-called cutting cone, was also rarely seen during the first year.
    In the medullary canal, ingrowth of fibrovascular reparative tissue surrounding necrotic cancellous trabeculae extended no more than a few millimeters with little or no new-bone formation. Internal repair was almost completely absent in eight specimens, involved about 5% of the graft in five, and involved about 10% of the graft in three.

    Second Year

    Twenty specimens were retrieved in the second year. During this period, the extent of osteoclastic surface resorption almost doubled, to approximately 40%. At the same time, the extent of surface apposition of new bone also doubled, to approximately 80%, filling in many of the resorption cavities seen in specimens retrieved earlier. However, in scattered areas of some specimens, an even pattern of repair was not seen. In these specimens, the external surface was pockmarked with unrepaired cavities ranging in size from several micrometers to a few millimeters. These cavities were filled with loosely arranged mesenchymal proliferations peppered with chronic inflammatory cells. Focal concentrations of inflammatory cells, foreign-body-type giant cells, or large histiocytes were only occasionally seen in these cavities. In some areas, such cavities were interspersed between areas of external repair by appositional new bone, while in others they were closely clumped together. Immediately beneath these areas of unrepaired surface erosion, no revascularization or internal repair was seen.
    At two years, somewhat deeper internal repair was directed by the haversian and Volkmann canals of the graft. Vessels and osteoclasts enlarged the diameter of the canals but rarely penetrated deeper than 5 mm from the cortical surfaces. Within the enlarged canals, osteoclastic resorption seldom extended beyond the peripheral cement lines of the surrounding osteons, and the interstitial lamellae beyond remained completely undisturbed by repair. In many of the enlarged canals, resorptive activity had ceased, and the walls of the canals were lined by seams of newly deposited living bone. Such repair occupied no more than 20% of the graft and was mostly confined to the peripheral aspects of the graft.
    At the osteotomy site, deeper penetration by revascularizing repair tissue was evident but lagged behind the repair from the surface in almost all specimens (Figs. 6-A and 6-B).
    In the medullary canal, the repair tissue extended up to 2 cm but only intermittently differentiated into osteoblasts with new bone laid on old necrotic trabeculae on the endosteal surface of the cortex. At the leading edge of this repair tissue, it frequently matured into dense fibrous tissue, impeding further reparative incursion.

    After the Second Year

    Twenty-two specimens retrieved more than two years after implantation were available for study: fifteen had been retrieved between two and five years after implantation and seven, between five and thirteen years.
    In the typical specimen, osteoclastic resorption was markedly decreased at the osteotomy site where union had occurred and on the external cortical surface. The thin layer of appositional bone covered a gradually greater proportion of the surface but had not increased in thickness. In contrast, revascularization and proliferation of osteoblasts repairing the excavated haversian canals and the Howship lacunae had increased. Typically, the osteoclastic erosion within an individual osteon had rarely reached the peripheral cement line or invaded the adjacent interstitial lamellae before it subsided and osteoblastic apposition superseded it (Figs. 7-A and 7-B). The cancellous trabeculae of the medullary canal and the endosteal surfaces of the graft remained unrepaired throughout the duration of the study.
    Although the extent of repair slowly progressed, it rarely penetrated >10 mm into the cortex. The variability in the extent of repair, however, was much greater in the group of specimens retrieved later than in the specimens retrieved after less than two years. In the typical specimen, the total extent of internal repair remained between 20% and 30%. However, a specimen retrieved at five years showed <5% repair, a specimen retrieved at thirteen years showed only 20% repair, and a specimen retrieved at eight years showed 70% repair.

    Soft-Tissue Attachment

    The interface between the allograft and the adjacent soft tissues was studied macroscopically in forty-three specimens. The soft tissues were attached to between 20% and 40% of the surface area in the specimens retrieved before six months, to between 50% and 80% of the surface area in specimens retrieved between six and twelve months, and to >80% of the surface area in those retrieved after twelve months.
    Labeling with tetracycline demonstrated the areas in which a thin seam of new external repair bone had been deposited upon the surface. Macroscopic and histological study of these areas showed that heavy strands of collagen ran from adjacent muscle, fascia, or ligament into the seam of new bone. Histological study under polarized light showed continuity of the collagen fibers as they extended from the soft tissues into the appositional new bone, but the collagen fibers did not cross the cement line that demarcated the living appositional new bone from the underlying acellular necrotic cortex of the allograft. In areas where the adherent soft tissues had been forcibly stripped from the graft, the tissues debonded along the demarcating cement line. In areas of substantial surface erosion in specimens that had been retrieved earlier, there was no bonding of the soft tissue as these areas were separated from the underlying bone by a film composed of fibrovascular repair tissue, scattered chronic inflammatory tissue, and an occasional foreign-body giant cell. When these erosion cavities had been filled in with appositional new bone in the specimens that had been retrieved later, the soft tissues became bonded as in the other areas.
    In the infected specimens, there was no seam of appositional new bone on the graft surface, and the surrounding edematous tissues were separated from the graft by a zone of chronic inflammatory granulation tissue with no soft-tissue attachment directly to the graft.

    Resorption of Allografts

    Eight specimens were retrieved because of resorption of substantial portions of the allograft. Consistent findings in these specimens were encasement of the graft by tissue composed of chronic inflammatory cells, histiocytes, and foreign-body giant cells in a background of proliferating mesenchymal cells. The acellular matrix was undergoing active osteoclastic resorption. There was no revascularization or internal repair of the remaining bone or any host-soft-tissue attachment. The specimens that had been retrieved later showed marked fragmentation. Bacterial cultures were consistently negative.
    A particularly interesting case was that of a fifteen-year-old girl who had reconstruction of the distal one-third of the femur with an osteoarticular allograft (Fig. 8-A). Plate fixation failed, and nonunion and fragmentation led to removal of the graft at fourteen months (Fig. 8-B). The defect was reconstructed with a second osteoarticular graft, which was securely united at eight months (Fig. 8-C). Gradually, over the next six months, there was marked resorption of the proximal portion of the graft (Fig. 8-D). The distal portion, including the joint, remained intact, and the patient had a well-functioning knee. In an effort to prevent a pathological fracture and to preserve the knee, a vascularized fibular graft was used to bypass the areas of greatest resorption. Curettings from the areas of resorption demonstrated findings similar to those described above. The vascular graft united to the host femur proximally and the allograft metaphysis distally, and, surprisingly, during the next twelve months the areas of resorption reossified and remained so for the following twelve years (Fig. 8-E).

    Allograft Fracture

    Thirteen specimens were retrieved because of fracture. In twelve specimens, the fracture was diaphyseal and was at the end of a plate (seven), at a screw-hole (four), or at the tip of a prosthetic stem (one). In one specimen, involving an entire femur, a fracture occurred midway between the tips of the stems of the total hip and total knee prostheses. The average interval to fracture was twenty-four months (range, thirteen to seventy-two months).
    In the specimens retrieved shortly following the fracture, histological examination of the site showed ingrowth of fibrovascular repair tissue into the gap with moderate osteoclastic resorption of the adjacent necrotic bone. There was no evidence of callus emanating from the graft fragments or unusual internal repair of the bone adjacent to the fracture line.
    Four specimens were retrieved several months after the fracture had occurred. One was retrieved because of malunion seven months after the fracture; one, after amputation because of a local recurrence eight months after the fracture; one, as a result of a biopsy of the site of a suspected tumor recurrence twelve months after the fracture; and one, at autopsy thirteen months after the fracture. The specimen retrieved at eight months had solid union of a diaphyseal fracture of the proximal part of the humerus by a large external callus. The specimen retrieved at autopsy was from a patient who had sustained a fracture of the tibia while receiving chemotherapy for pulmonary metastases. The fracture had been treated with immobilization in a cast for seven months, and it had healed with an external callus at the time of autopsy thirteen months later.
    The capacity of repairing allografts to heal was illustrated by the case of an eleven-year-old girl who had reconstruction of the distal part of the femur with a 27-cm osteoarticular graft after resection of a stage-IIB osteosarcoma2 (Fig. 9-A). Four years later, she sustained a fracture through the metaphysis of the allograft (Fig. 9-B), which was treated with a plaster cast and healed with a substantial malunion. Because of concomitant degenerative changes in the articular cartilage, the distal 15 cm of the allograft, including the site of the malunion, was resected, preserving the more proximal 12 cm of the original allograft. This defect was reconstructed with a composite allograft-total knee prosthesis fixed by a long intramedullary stem extending well above the junction of the original allograft and the second allograft. Supplementary autogenous iliac-crest grafts were secured about the junction with wires. The junction had united at twelve months (Fig. 9-C), and the reconstruction functioned well for the ensuing seven years (Fig. 9-D). Histological study of the resected malunion site in the original graft showed internal repair of almost 60% of the cortices at the fracture site compared with <20% in the remainder of the graft.
    The common finding in these specimens was an unusual amount of internal repair in the region about the fracture. Spatial maps showed extensive revascularization in both the cortices and the trabeculae, extending several centimeters on either side of the fracture, paralleling the extent of the external callus.

    Allograft-Cement Interface

    Eight allograft-prosthesis composites were retrieved. Macrosections of five of these specimens, retrieved at intervals of seven months to eight years after implantation, were available for study. There was no radiographic evidence of resorption of the allograft about the cement. The interface between the allograft and the cement was grossly secure in all specimens. On histological examination, none of the specimens demonstrated revascularization of the allograft adjacent to the cement, ingrowth of tissue between the cement and adjacent allograft, or disruption of the allograft architecture.

    Frozen Articular Cartilage

    The macroscopic and histological features of the articular cartilage were studied in twenty-eight preserved osteoarticular allografts. Twenty-four specimens were totally devoid of chondrocytes in the lacunae of the persisting articular cartilage. Four specimens, all retrieved within eight months after implantation, had occasional lacunae with remnants of cells that appeared to represent mummification of chondrocytes during the freezing and storage process.
    The five specimens retrieved prior to one year (average, seven months) showed no change radiographically either in the apparent thickness of the articular cartilage or in the underlying subchondral bone. All had surface irregularities of varying severity with fibrillation of the superficial surface of the cartilage, tangential clefts with flap separations, erosions, and, occasionally, perpendicular clefts reaching as deep as the tidemark (Fig. 10).
    Eight specimens were retrieved between one and two years (average, seventeen months) after implantation. Again, radiographically, the thickness of the articular cartilage showed little or no narrowing, and the subchondral bone appeared to be intact with no evidence of repair or fragmentation. The most striking change was the extent of a pannus of fibrovascular repair tissue covering some or all of the articular surfaces. The pannus emanated from the periphery of the joint and extended to varying degrees until the entire surface was covered (Fig. 11). As the pannus first developed, it covered the free surface, actually increasing the thickness of the cartilage as seen radiographically, but, with time, the articular cartilage beneath the pannus was resorbed, resulting in a combination pannus-cartilage remnant that appeared to be thinner radiographically. As the pannus covered the surface, it frequently extended down into the clefts in the cartilage in an apparent attempt to repair them. As the pannus thickened, it occasionally contained nodules of fibrocartilage derived from the proliferating host mesenchymal cells within it.
    Nine specimens were retrieved between twenty-four and sixty months (average, forty-four months) after implantation. The radiographs showed moderate thinning of the pannus-cartilage remnant and increasing radiolucency of the cancellous bone beneath the subchondral plate. In many of these specimens, there were varying degrees of subchondral fragmentation and deformation. The macroscopic and histological pattern was one of increasing resorption of the remaining articular cartilage, which, in some specimens, reached to the tidemark but rarely extended across it to the subchondral plate.
    All of these changes progressed with time and were more advanced in specimens from the lower extremity than in those from the upper extremity.
    Four specimens were retrieved between five and ten years (average, seven years) after implantation, and two were retrieved after more than ten years. These specimens had considerable variation in the amount of cartilage destruction and the extent of subchondral fragmentation or deformation.

    Fresh Articular Cartilage

    One fresh osteoarticular graft was a hemiarticular femoral condyle and distal metaphysis harvested from a heart-beating donor and implanted following resection of a presumed stage-3 aneurysmal bone cyst3. Postoperatively, the patient was managed with immunosuppression with prednisone and Imuran (azathioprine) on the schedule then used for organ recipients. The graft was fixed with multiple screws and was united at four months. The patient regained 90° of motion and began bearing weight at seven months. The patient died of metastatic giant-cell tumor, and the specimen was retrieved at autopsy thirteen months after implantation. The articular cartilage was well preserved with no degenerative changes. Histological examination showed a composition of viable chondrocytes without fibrillation or clefts and a distinct intact tidemark (Figs. 12-A and 12-B).
    The other freshly transplanted osteoarticular graft, a fetal femoral head that was used to replace a metacarpal head destroyed by psoriatic arthropathy, was retrieved at three months during a ray resection. No immunosuppression had been used. The specimen was populated with viable-appearing chondrocytes but was surrounded by a zone of inflammatory tissue.
    The findings in our initial report1 were supported and confirmed by the observations in the present study, which involved a much larger group of specimens obtained over a longer period of time. However, the present group of specimens, particularly those retrieved at longer intervals, had much greater heterogeneity, which provided additional insights that suggest that the interaction between the host and allograft is influenced by several factors.
    Since more than half of the specimens had been retrieved because of a complication associated with the allograft that may have altered the usual course of repair, the findings in the twenty-four specimens (33%) retrieved because of amputation or at autopsy may be more representative of the reparative mechanisms than the findings in the group as a whole. However, when the two groups were analyzed with respect to the timing and extent of the repair, no important differences were noted except in the specimens removed because of early infection (five) or resorption (eight). In those specimens, both external and internal repair as well as soft-tissue adherence were either markedly diminished compared with the findings in the specimens without graft complications or they were absent. Overall, however, the majority of the specimens retrieved because of graft complications had findings similar to those in the specimens without such complications.
    Accurate and intimate contact between host and allograft cortices appears to promote and accelerate union, although healing was also observed when gaps of up to 4 mm were present at securely immobilized junctions. The degree of contact and the security of fixation appeared to influence the size and extent of the external callus and its maturation into haversian bone. However, in no instance was there evidence of primary union, even when there was the most intimate contact; this was due, at least in part, to the inability of the host fibrovascular repair tissue to produce cutting cones or to effectively penetrate the allograft haversian systems. The persistent failure of the host bone that filled the gaps to remodel along stress lines and the fact that the junction of this bone with the allograft was identified as the site of induced debonding indicate that an allograft-host junction may remain a potential site for subsequent failure for an extended period of time. Whether such remodeling would be induced by removal of the internal fixation devices remains speculative, and, in view of this, the additional support of fixation devices may well be advantageous in this regard. It would also appear to be desirable to quantitate the extent of this weakness with biomechanical studies so that effective means of prevention might be developed.
    The contrast between histologically united cortical-cortical junctions and the persistence of radiolucency in the intercortical gaps reaffirms the prudence of postponing operative intervention because of presumed nonunion for at least one year until maturation of the bone is reflected radiographically. This is especially true in patients who receive postoperative chemotherapy. Cortical-cortical union appeared to be substantially enhanced when autogenous bone grafts had been added, with active callus formation extending onto the external surface of the graft. The osteoinductive properties of autogenous grafts are likely responsible for the increase in internal repair of the adjacent cortex of the allograft4, which raises the hypothesis that the use of more recently developed osteoinductive substances may be similarly useful.
    The antiblastic toxicity of chemotherapeutic drugs is known to have an inhibitory effect on allograft union and repair in animals5. This is reflected histologically by diminished osteoclastic resorption and decreased new-bone formation resulting from suppression of osteoblastic activity and abolition of mesenchymal proliferation. In our group of human allograft specimens, preoperative chemotherapy was associated with retarded union. Union was rare before one year, and maturation of callus required more than one year. These observations suggest that, in humans, the deleterious effect of chemotherapy is a reversible process, with restoration of a biological environment favorable to allograft healing occurring at the end of the chemotherapy period. In contrast, a specimen from a patient who received substantial amounts of postoperative radiation therapy showed no signs of union three years after implantation, suggesting that radiation damage permanently jeopardizes allograft incorporation.
    Repair of the grafts occurred in two fashions—surface (or external) repair and internal repair. Surface repair was accomplished by the deposition of a thin seam of appositional host bone, beginning within the first three to six months, on the unresorbed surface of the graft that served as the anchor for the attachment of host soft tissues. However, the demarcating cement line was the site of debonding when the soft tissues were forcibly pulled away. After maturation into lamellar bone, usually by one year, no additional thickening of the seam occurred. On average, by one year such deposition covered approximately one-half of the exposed cortical surfaces. The remaining cortical surfaces underwent superficial resorption in a random fashion. These surfaces became pockmarked with Howship lacunae, only a few of which contained osteoclasts at any given time. Such surface resorption removed only a superficial millimeter or two and, with cessation, was filled in by appositional new bone so that by two years almost the entire surface of the graft was covered by a seam of viable lamellar bone. Although too thin to be apparent on either conventional radiography or computed tomography, it was evidenced by a correspondingly thin rim of increased activity on isotope scanning.
    Internal repair began with invasion of the surface stoma of Volkmann and haversian canals by fibrovascular host tissue that enlarged the haversian canals by osteoclastic resorption and then rebuilt them with appositional new bone. The pattern of such revascularization was random, and the pace was slow, seldom exceeding more than a few millimeters per year. A repaired osteon was often seen side by side with one that was totally ignored by the reparative tissues. Such internal repair rarely occurred from the cut ends of the graft, proceeding longitudinally into the old haversian canals, and almost never by cutting cones at either the surface or the osteotomy sites. The domination of resorptive activity during the first two years suggests that the second year after implantation is the critical period of cortical weakening of massive allografts. The extent and pace of internal repair in cortical bone were so limited that they did not alter the radiographic density of the graft with areas of radiolucency or bind enough isotope to produce visible increases in uptake on isotope scanning. This general picture was common to all repairing grafts for two to three years. Specimens retrieved after three years had a much wider variation in the extent of cortical repair. In one specimen, cortical repair virtually ceased; in some, it sputtered along; and in a few, it progressed steadily until virtually the entire graft was repaired.
    In the present study, there was a strong histological suggestion of an immunological mechanism in the specimens that had undergone resorption without repair. Each one had aggressive resorption of cortical bone by osteoclasts crossing the peripheral cement line about the osteons and resorbing the interstitial lamellae, a pattern not seen in autogenous cortical graft repair6,7 and one that is common in xenograft resorption8,9.
    Areas of unusually extensive repair in the cortices were seen under several diverse circumstances: at sites of union with supplementary conventional autogenous cancellous grafts, at sites of union with supplementary autogenous vascularized grafts, about the sites of healing allograft fractures, and at sites of involvement of the allograft by recurrent neoplastic tissue. There was no apparent reason for this phenomenon, and the mechanism appeared to be a localized nonspecific acceleration of the usual reparative response.
    As suggested by the histological findings and as reported by others10-12, in our study allograft fractures occurred more frequently during the second year after implantation, a reflection of the domination of the resorptive pattern during this phase of the remodeling process. Diaphyseal fractures occurred in areas of stress concentration about the ends of fixation devices or where the cortex had been perforated by screws. In two patients in whom autogenous cancellous grafts had been placed at the osteotomy site, a fracture occurred just above the region of accelerated internal repair adjacent to the autogenous grafts. Similarly, subchondral fractures occurred through the metaphyseal cortex where fibrovascular repair tissue and osteoclastic resorption had extended into the subchondral bone.
    The longer periods of observation of the interface between allograft bone and bone cement in these specimens confirmed the earlier observation1 that there is little or no change in the microarchitecture at the bone-cement interface and that the usual lack of extensive cortical revascularization is advantageous in this regard. Given these observations, it seems prudent to use intramedullary fixation rather than plates and screws whenever practical, to use bone cement to firmly anchor fixation devices and prosthetic stems in composite reconstructions, and to leave fixation devices in situ indefinitely unless device-related disability mandates removal.
    The survival of chondrocytes in the articular cartilage of frozen allografts treated with cryoprotectants has been a controversial subject1,4,13-16. Delloye et al. cultivated frozen cartilage treated with dimethyl sulfoxide (DMSO) and observed cell growth with the morphological appearance of living chondrocytes but did not observe any differences with the control group (allografts frozen without cryoprotectant)13. Malinin et al. reported cell growth after cultivating cartilage from an articular allograft stored in liquid nitrogen, although they did not establish the cartilaginous nature of these cells14. Schachar and McGann studied the viability of isolated chondrocytes both in vitro and in animal models16. They asserted that the persistent viability of cartilaginous cells was optimized by the use of DMSO as a cryoprotectant and by utilizing slow freezing and rapid thawing of the graft. However, we observed only scattered remnants of nucleated chondrocytes with cytomorphological features of viability in four of twenty-eight osteoarticular specimens. Quite clearly, the majority of human chondrocytes stored by conventional banking techniques do not survive despite the use of cryoprotectants. In contrast, two fresh specimens had clear histological evidence of chondrocyte survival, confirming the observation of Oakeshott et al. that fresh, unfrozen allografts had substantial chondrocyte survival17. Their small allografts had little or no inflammatory response. In our study of larger specimens, a marked inflammatory response had occurred in the fresh specimen from the patient who had not had immunosuppression, whereas evidence of widespread chondrocyte survival with a minimal inflammatory response was seen in the specimen retrieved at thirteen months from the patient who had had immunosuppression. These observations suggest that fresh, unfrozen chondrocytes survive, but patients receiving a large osteochondral graft may require immunosuppression for the chondrocytes to remain viable.
    In the osteoarticular specimens retrieved later (after two years or more), formation of the fibrovascular pannus took two forms. In some specimens, it formed a thin covering over the necrotic cartilage protruding into and filling clefts and it appeared to be primarily reparative in nature. In other specimens, the pannus was much thicker, was more cellular, and consisted primarily of chronic inflammatory cells, and it appeared to be primarily inflammatory in nature, again suggesting that these differences are based on histocompatibility. It may be that when early technical disruption exposes previously immunologically sequestrated chondrocytes to the host immune system, those with a major immunological mismatch incite an inflammatory response whereas those with a minor mismatch evoke a lesser immunological stimulus to form a reparative response. This hypothesis is also suggested by the observation that, without exception, every specimen had mechanically induced changes in the articular cartilage, whereas the formation of pannus was an inconstant and later event.
    Abnormal anatomical and dimensional matching of the articular surfaces and capsuloligamentous instability appeared to play a role in the rapidity and degree of articular cartilage degeneration. In the specimens retrieved earlier (after less than two years), anatomical mismatching and/or instability appeared to be responsible for irregular distribution of mechanical forces on the articular surface, which created stress-raising areas in which degeneration with fibrillation and tangential flap separation, representing the initial phase of osteoarthritis, took place. The hypothesis that there is a mechanical role in the initiation of degenerative changes is reinforced by the higher prevalence of such changes in the weight-bearing knee than in the non-weight-bearing shoulder.
    It was of considerable interest that for the first one to two years, despite substantial clinical activity, the necrotic articular cartilage functioned well and appeared to be normal radiographically. In fact, on the radiographs of many specimens, it was not possible to distinguish between persisting articular cartilage with a thin pannus and major cartilage destruction and thick inflammatory pannus. Only when substantial subchondral bone resorption and fragmentation occurred were clinical and radiographic signs of substantial narrowing of articular cartilage apparent. In some instances this occurred at one year, and in others it did not occur for more than five years.
    Enneking WF, and Mindell ER: Observations on massive retrieved human allografts. J Bone Joint Surg Am,1991.73: 1123-42, 731123  1991  [PubMed]
     
    Enneking WF; Spanier SS; and Goodman MA: Current concepts review. The surgical staging of musculoskeletal sarcoma. J Bone Joint Surg Am,1980.62: 1027-30, 621027  1980  [PubMed]
     
    Enneking WF: A system of staging musculoskeletal neoplasms. Clin Orthop,1986.204: 9-24, 2049  1986  [PubMed]
     
    Urist MR: Bone: formation by autoinduction. Science,1965.150: 893-9, 150893  1965  [PubMed]
     
    Friedlaender GE; Tross RB; Doganis AC; Kirkwood JM; and Baron R: Effects of chemotherapeutic agents on bone. I. Short-term methotrexate and doxorubicin (Adriamycin) treatment in a rat model. J Bone Joint Surg Am,1984.66: 602-7, 66602  1984  [PubMed]
     
    Burchardt H; Busbee GA 3rd; and Enneking WF: Repair of experimental autologous grafts of cortical bone. J Bone Joint Surg Am,1975.57: 814-9, 57814  1975  [PubMed]
     
    Enneking WF, and Morris JL: Human autologous cortical bone transplants. Clin Orthop,1972.87: 28-35, 8728  1972  [PubMed]
     
    Goldberg VM; Powell A; Shaffer JW; Zika J; Bos GD; and Heiple KG: Bone grafting: role of histocompatibility in transplantation. J Orthop Res,1985.3: 389-404, 3389  1985  [PubMed]
     
    Kirkeby OJ; Nordsletten L; and Skjeldal S: Healing of cortical bone grafts in athymic rats. Acta Orthop Scand,1992.63: 318-22, 63318  1992  [PubMed]
     
    Berrey BH Jr; Lord CF; Gebhardt MC; and Mankin HJ: Fractures of allografts. Frequency, treatment, and end-results. J Bone Joint Surg Am,1990.72: 825-33, 72825  1990  [PubMed]
     
    Thompson RC Jr; Pickvance EA; and Garry D: Fractures in large-segment allografts. J Bone Joint Surg Am,1993.75: 1663-73, 751663  1993  [PubMed]
     
    Zehr RJ, Enneking WF, Heare T, Liang TS. Fractures in large structural allografts. In: Brown KLB, editor. Complications of limb salvage. Montreal: International Symposium on Limb Salvage; 1991. p 3-8. 
     
    Delloye C; De Halleux J; Cornu O; Wegmann E; Buccafusca GC; and Gigi J: Organizational and investigational aspects of bone banking in Belgium. Acta Orthop Belg,1991.57(Suppl 2): 27-34, 57(Suppl 2)27  1991  [PubMed]
     
    Malinin TI; Martinez OV; and Brown MD: Banking of massive osteoarticular and intercalary bone allografts—12 years’ experience. Clin Orthop,1985.197: 44-57, 19744  1985  [PubMed]
     
    Mankin HJ; Doppelt S; and Tomford W: Clinical experience with allograft transplantation. The first ten years. Clin Orthop,1983.174: 69-86, 17469  1983  [PubMed]
     
    Schachar NS, McGann LE. Cryopreservation of articular cartilage. In: Friedlaender GE, Goldberg VM, editors. Bone and cartilage allografts. American Academy of Orthopaedic Surgeons Symposium. 1989. p 221-30. 
     
    Oakeshott RD; Farine I; Pritzker KP; Langer F; and Gross AE: A clinical and histologic analysis of failed fresh osteochondral allografts. Clin Orthop,1988.233: 283-94, 233283  1988  [PubMed]
     

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    +Fig. 1:Graph showing the number of specimens retrieved per year.
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    +Fig. 2-A:Figs. 2-A and 2-B The histological features of cortical-cortical union. Fig. 2-A A low-power photomicrograph showing the junction between the acellular cortex of an allograft (AG) and the host bone (HB) filling the gap (hematoxylin and eosin, 20). The allograft was retrieved at five years. The site of union is marked by a distinct cement line (CL). Only a single haversian canal (HC) has been revascularized.
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    +Fig. 2-B:Figs. 2-A and 2-B The histological features of cortical-cortical union. Fig. 2-B A low-power photomicrograph showing the junction between the cortex of an allograft (AG) and the host bone (HB) filling the gap (hematoxylin and eosin, 20). The allograft was retrieved at eleven years. The site of union is still marked by a distinct cement line (CL). Revascularization has penetrated two haversian canals (HC).
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    +Fig. 3-A:Figs. 3-A and 3-B The site of failure at forty-two months. Fig. 3-A A photomicrograph showing the failure site between the cortex of an allograft (AG) and the host cortex (HE), which has extended through the mature external callus (EC) that united the two cortices (hematoxylin and eosin, 4).
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    +Fig. 3-B:Figs. 3-A and 3-B The site of failure at forty-two months. Fig. 3-B A photomicrograph showing the failure site between the allograft (AG) and the nonoriented host bone (HB) at the demarcating cement line (CL) (hematoxylin and eosin, 40).
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    +Fig. 4-A:Figs. 4-A and 4-B Union by endosteal callus. Fig. 4-A Anteroposterior radiograph of a proximal tibial osteoarticular allograft immediately before retrieval at seventeen months. No external callus is visible, and there is no radiolucent line at the junction (J) of the allograft and the more osteopenic host tibia.
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    +Fig. 4-B:A low-power photomicrograph of the allograft cortices (AG) and the more osteopenic host cortices (HC) (hematoxylin and eosin, 10). The upper junction is united by internal endosteal callus (IC), while the lower junction remains ununited despite the well-formed internal callus (IC).
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    +Fig. 5:A photomicrograph of the surface of an allograft retrieved eleven months after implantation, showing surface repair of the allograft cortical bone (hematoxylin and eosin, 120). The host soft tissues (HST) are apposed to the surface of the allograft cortex. A bud of reparative tissue entered the surface stoma of a haversian canal (HC), enlarging it by osteoclastic resorption. A thin seam of bone (HB) has been laid down on the surface of the allograft and is demarcated from it by a distinct cement line (CL).
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    +Fig. 6-A:Figs. 6-A and 6-B Revascularization at the osteotomy site. Fig. 6-A A photomicrograph showing the cortical osteotomy site of an allograft retrieved nineteen months after implantation (hematoxylin and eosin, ¥60). The three arrows in the center of the figure indicate the site of union between the host bone (HB) and the allograft (AG). Reparative tissue has penetrated into the graft in two areas (RT), enlarging the haversian canals (HC), but several canals show no evidence of repair.
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    +Fig. 6-B:A photomicrograph showing the cortical osteotomy site of an allograft retrieved twenty-five months after implantation (hematoxylin and eosin, ¥90). The viable host bone (HB) is united to the necrotic allograft (AG). The six arrows at the left of the figure indicate a previously enlarged haversian canal, filled with repair bone (RB). A second haversian canal contains reparative tissue (RT). Other haversian canals (HC) remain unrepaired.
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    +Fig. 7-A:Figs. 7-A and 7-B Internal repair. Fig. 7-A A low-power photomicrograph of a repairing allograft retrieved at thirty months (hematoxylin and eosin, 20). The host muscle (HM) is separated from the repairing allograft (AG) by a thin zone of proliferating host fibrovascular repair tissue. The haversian canals of the superficial osteons (SO) are largely excavated, leaving the interstitial lamellae (IL) unrepaired, while the deeper osteons are not yet revascularized and their haversian canals (HC) are not yet excavated.
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    +Fig. 7-B:A photomicrograph of osteons being reformed in a cortical allograft retrieved at twenty-six months (hematoxylin and eosin, 90). Osteoclastic resorption has almost reached the cement line (arrows) bounding the osteon on the left, while on the right excavation has reached the bounding cement line (arrows) about the osteon, the osteoclasts have disappeared, and osteoblastic rebuilding (OB) has begun.
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    +Fig. 8-A:Figs. 8-A through 8-E Reossification of allograft resorption. Fig. 8-A Anteroposterior radiograph made after implantation of the initial osteoarticular allograft.
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    +Fig. 8-B:Figs. 8-A through 8-E Reossification of allograft resorption. Fig. 8-B Anteroposterior radiograph made fourteen months later.
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    +Fig. 8-C:Fig. 8-C Anteroposterior radiograph made after implantation of the second osteoarticular allograft.
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    +Fig. 8-D:Fig. 8-D Anteroposterior radiograph made six months later.
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    +Fig. 8-E:Fig. 8-E Anteroposterior radiograph made twelve years after curettage and insertion of a supplementary vascular autogenous fibular graft.
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    +Fig. 9-A:Figs. 9-A through 9-D Union of an allograft-allograft junction. Fig. 9-A Postoperative anteroposterior radiograph of a 27-cm femoral osteoarticular allograft. The arrow indicates the allograft-host junction (J).
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    +Fig. 9-B:Figs. 9-A through 9-D Union of an allograft-allograft junction. Fig. 9-B Anteroposterior radiograph made four years postoperatively, at the time of distal allograft fracture. The allograft-host junction is well healed (arrow).
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    +Fig. 9-C:Fig. 9-C Anteroposterior radiograph made one year after a second reconstruction with a 15-cm allograft-prosthesis composite. The arrow indicates the junction of the first and second allografts.
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    +Fig. 9-D:Fig. 9-D Magnified anteroposterior radiograph of the junction of the first and second allografts, made seven years after the second reconstruction.
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    +Fig. 10:Early degenerative changes in articular cartilage. A photomicrograph of the articular surface of a proximal tibial osteoarticular allograft retrieved at eleven months (hematoxylin and eosin, 60). The cartilage is acellular and has severe degenerative changes, although the tidemark (TM) remains intact. The underlying subchondral marrow spaces have been revascularized, although there has been little repair of bone.
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    +Fig. 11:Pannus formation. A photomicrograph of the posterior aspect of the articulating surface of a femoral condyle from a distal femoral osteoarticular allograft retrieved at seventeen months (hematoxylin and eosin, 40). The articular cartilage (C) is acellular. A thick pannus of fibrovascular tissue (FVT) covers the cartilage. A tongue of more cellular tissue invades the cartilage at its periphery, having crossed both the tidemark (TM) and the osteochondral cement line (CL).
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    +Fig. 12-A:Figs. 12-A and 12-B Survival of fresh allograft chondrocytes. Fig. 12-A A photomicrograph of a fresh osteoarticular distal femoral allograft retrieved at thirteen months (hematoxylin and eosin, 5). The architecture of both the articular cartilage and the underlying femoral condyle is well preserved.
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    +Fig. 12-B:A photomicrograph of a representative field of the articular cartilage shows the normal architecture of adult articular cartilage (hematoxylin and eosin, 90). The vast majority of the chondrocytes appear viable, with only small patches of acellular matrix. There is no evidence of an inflammatory response.
    Enneking WF, and Mindell ER: Observations on massive retrieved human allografts. J Bone Joint Surg Am,1991.73: 1123-42, 731123  1991  [PubMed]
     
    Enneking WF; Spanier SS; and Goodman MA: Current concepts review. The surgical staging of musculoskeletal sarcoma. J Bone Joint Surg Am,1980.62: 1027-30, 621027  1980  [PubMed]
     
    Enneking WF: A system of staging musculoskeletal neoplasms. Clin Orthop,1986.204: 9-24, 2049  1986  [PubMed]
     
    Urist MR: Bone: formation by autoinduction. Science,1965.150: 893-9, 150893  1965  [PubMed]
     
    Friedlaender GE; Tross RB; Doganis AC; Kirkwood JM; and Baron R: Effects of chemotherapeutic agents on bone. I. Short-term methotrexate and doxorubicin (Adriamycin) treatment in a rat model. J Bone Joint Surg Am,1984.66: 602-7, 66602  1984  [PubMed]
     
    Burchardt H; Busbee GA 3rd; and Enneking WF: Repair of experimental autologous grafts of cortical bone. J Bone Joint Surg Am,1975.57: 814-9, 57814  1975  [PubMed]
     
    Enneking WF, and Morris JL: Human autologous cortical bone transplants. Clin Orthop,1972.87: 28-35, 8728  1972  [PubMed]
     
    Goldberg VM; Powell A; Shaffer JW; Zika J; Bos GD; and Heiple KG: Bone grafting: role of histocompatibility in transplantation. J Orthop Res,1985.3: 389-404, 3389  1985  [PubMed]
     
    Kirkeby OJ; Nordsletten L; and Skjeldal S: Healing of cortical bone grafts in athymic rats. Acta Orthop Scand,1992.63: 318-22, 63318  1992  [PubMed]
     
    Berrey BH Jr; Lord CF; Gebhardt MC; and Mankin HJ: Fractures of allografts. Frequency, treatment, and end-results. J Bone Joint Surg Am,1990.72: 825-33, 72825  1990  [PubMed]
     
    Thompson RC Jr; Pickvance EA; and Garry D: Fractures in large-segment allografts. J Bone Joint Surg Am,1993.75: 1663-73, 751663  1993  [PubMed]
     
    Zehr RJ, Enneking WF, Heare T, Liang TS. Fractures in large structural allografts. In: Brown KLB, editor. Complications of limb salvage. Montreal: International Symposium on Limb Salvage; 1991. p 3-8. 
     
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